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Encyclopedia > Atlantic herring

Atlantic herring

Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Actinopterygii
Order: Clupeiformes
Family: Clupeidae
Genus: Clupea
Species: C. harengus
Clupea harengus
Linnaeus, 1758

Atlantic herring Clupea harengus is the one of the most abundant species of fish on the planet. They can be found on both sides of the Atlantic Ocean congregating together in large schools or (swarms). They can grow up to 45 centimeters (approximately 18 inches) in length and weigh more than half a kilogram. They feed on copepods, krill and small fish, their natural predators are seals, whales, cod and other larger fish. Image File history File links atlantic herring image uwe kils This work is copyrighted. ... Scientific classification or biological classification is how biologists group and categorize extinct and living species of organisms (as opposed to folk taxonomy). ... Phyla Subregnum Parazoa Porifera (sponges) Subregnum Agnotozoa Placozoa (trichoplax) Orthonectida (orthonectids) Rhombozoa (dicyemids) Subregnum Eumetazoa Radiata (unranked) (radial symmetry) Ctenophora (comb jellies) Cnidaria (coral, jellyfish, anemones) Bilateria (unranked) (bilateral symmetry) Acoelomorpha (basal) Orthonectida (parasitic to flatworms, echinoderms, etc. ... Typical Classes Subphylum Urochordata - Tunicates Ascidiacea Thaliacea Larvacea Subphylum Cephalochordata - Lancelets Subphylum Myxini - Hagfishes Subphylum Vertebrata - Vertebrates Petromyzontida - Lampreys Placodermi (extinct) Chondrichthyes - Cartilaginous fishes Acanthodii (extinct) Actinopterygii - Ray-finned fishes Actinistia - Coelacanths Dipnoi - Lungfishes Amphibia - Amphibians Reptilia - Reptiles Aves - Birds Mammalia - Mammals Chordates (phylum Chordata) include the vertebrates, together with... Orders See text The Actinopterygii are the ray-finned fish. ... Families Denticipitidae (denticle herring) Engraulidae (anchovies) Pristigasteridae (pristigasterids) Chirocentridae (wolf herring) Clupeidae (herrings) Clupeiformes is the order of ray-finned fish that includes the herring family, Clupeidae, and the anchovy family, Engraulidae. ... Genera Subfamily Dussumieriinae (round herrings)     Dayella     Dussumieria     Etrumeus     Gilchristella     Jenkinsia     Luisiella     Sauvagella     Spratelloides     Spratellomorpha Subfamily Clupeinae     Amblygaster     Clupea     Clupeonella     Escualosa     Harengula     Herklotsichthys     Lile     Opisthonema Subfamily Alosinae (shads, menhadens)     Alosa     Brevoortia     Caspialosa     Ethmalosa     Ethmidium     Gudusia     Hilsa     Tenualosa Subfamily Pellonulinae     Knightia     Clupeichthys     Clupeoides     Congothrissa     Corica     Cynothrissa     Ehirava     Hyperlophus     Laeviscutella     Limnothrissa     Microthrissa     Odaxothrissa... Clupea may refer to: Clupea, the genus of herrings. ... In biology, binomial nomenclature is the formal method of naming species. ... Carolus Linnaeus Carl Linnaeus, also known after his ennoblement as , (May 23, 1707 – January 10, 1778), was a Swedish botanist, physician and zoologist[1] who laid the foundations for the modern scheme of nomenclature. ... Atlantic herring, Clupea harengus: one of the most abundant species of fish in the world. ... School of juvenile herring - many fish have the opercula wide open for ram feeding and you can see the red gills The term swarm (schooling or swarming) is applied to fish, birds and insects and describes a behavior of an aggregation (school) of animals of similar size and body orientation... To help compare different orders of magnitude this page lists lengths between 10-1 m and 1 m (10 cm and 1 m). ... The international prototype, made of platinum-iridium, which is kept at the BIPM under conditions specified by the 1st CGPM in 1889. ... Orders Calanoida Cyclopoida Gelyelloida Harpacticoida Misophrioida Monstrilloida Mormonilloida Platycopioida Poecilostomatoida Siphonostomatoida Copepods are a group of small crustaceans found in the sea and nearly every freshwater habitat. ... Families Euphausiidae Euphausia Dana, 1852 Meganyctiphanes Holt and W. M. Tattersall, 1905 Nematobrachion Calman, 1905 Nematoscelis G. O. Sars, 1883 Nyctiphanes G. O. Sars, 1883 Pseudeuphausia Hansen, 1910 Stylocheiron G. O. Sars, 1883 Tessarabrachion Hansen, 1911 Thysanoessa Brandt, 1851 Thysanopoda Latreille, 1831 Bentheuphausiidae Bentheuphausia amblyops Krill are shrimp-like marine... subfamilies Otariidae Phocidae Odobenidae Pinnipeds are large marine mammals belonging to the Pinnipedia, a family (sometimes a suborder or superfamily, depending on the classification scheme) of the order Carnivora. ... Whales are the largest species of exclusively aquatic mammals, members of the order Cetacea, which also includes dolphins and porpoises. ... Species Gadus morhua Gadus macrocephalus Gadus ogac Cod surfacing This article is about codfish; for other meanings, see COD. Cod is the common name for the genus Gadus of fish, belonging to the family Gadidae, and is also used in the common name of a variety of other fishes. ...


The Atlantic herring fishery has long been an important part of the economy of New England and the Canadian Maritime provinces, this is because the fish congregate relatively near to the coast in massive schools, notably in the cold waters of the semi-enclosed Gulf of Maine and Gulf of St. Lawrence. North Atlantic herring schools have been measured up to 4 cubic kilometers in size, containing an estimated 4 billion fish. Flag of New England The states of New England are Connecticut, Maine, Massachusetts, New Hampshire, Rhode Island, and Vermont. ... The Maritimes or Maritime provinces are a region of Canada on the Atlantic coast, consisting of New Brunswick, Nova Scotia and Prince Edward Island. ... Gulf of Maine The Gulf of Maine is a large gulf of the Atlantic Ocean on the northeastern coast of North America, roughly between Cape Cod in Massachusetts on the south and Cape Sable Island on the southern tip of Nova Scotia to the northeast. ... Gulf of Saint Lawrence, the worlds hugest estuary, is the outlet of North Americas Great Lakes via the Saint Lawrence River into the Atlantic Ocean. ... (Redirected from 1 E9 m3) To help compare different orders of magnitudes this page lists volumes between 1 and 10 cubic kilometres (109 to 1010 m³). See also volumes or capacities of other orders of magnitude. ...

Contents


Morphology

Atlantic herring have an elongated body that is fairly slender, a belly that is rounded (compared with that of a sprat, Sprattus sprattus), they also have no adipose fin this feature distinguishes herrings from the Family of salmon. The Atlantic herring are distinguished from other herring (there are close to 200 species in the family clupeidae) by their relativley small size, they have scutes without a prominent keel and they have a pelvic fin that is located behind the dorsal fin, the dorsal fin is located midway along their body. Atlantic herring can be further identified from that of other herring by their cluster of small teeth that are arranged in the shape of an oval at the roof of its mouth. This feature is particular to Atlantic herring. Sprat is a name given to a number of small, herring-like, marine fish in the family Clupeidae, notably in the genera Clupeonella and Sprattus. ... Binomial name Sprattus sprattus (Linnaeus, 1758) The European sprat is a small, herring-like, marine fish. ... Fish anatomy is primarily governed by the physical characteristics of water, which is much denser than air, holds a relatively small amount of dissolved oxygen, and absorbs light more than does air. ... Species Clupea alba Clupea bentincki Clupea caspiopontica Clupea chrysotaenia Clupea elongata Clupea halec Clupea harengus Clupea inermis Clupea leachii Clupea lineolata Clupea minima Clupea mirabilis Clupea pallasii Clupea sardinacaroli Clupea sulcata Herrings are small oily fish of the genus Clupea found in the temperate, shallow waters of the North Atlantic... The Chinook or King Salmon is the largest salmon in North America and can grow up to 58 long and 125 pounds. ... A scute (Latin scutum, shield) is a horny, chitinous, or bony external plate or scale, as on the shell of a turtle or the underside of a snake. ... A fer is a large beam around which the hull of a ship is built. ... Human male pelvis, viewed from front Human female pelvis, viewed from front The pelvis is the bony structure located at the base of the spine (properly known as the caudal end). ... The word dorsal can refer to many different things. ...


Ecological importance

Herring-like fish are the most important fish group on the planet, Clupea harengus the most frequent fish (Guinness Book of Records). They are the dominant converter of the enormous production of zooplankton, utilizing the biomass of copepods, mysids and krill in the pelagial. They are on the other side a central prey item for higher trophic levels. The reasons for its success is still enigmatic; one speculation is attributing their dominance to the outstanding way of living in huge, extremely fast cruising schools. Suresh Joachim, minutes away from breaking the ironing world record at 55 hours and 5 minutes, at Shoppers World, Brampton. ... Photomontage of plankton organisms Plankton is the aggregate community of weakly swimming but mostly drifting small organisms that inhabit the water column of the ocean, seas, and bodies of freshwater. ... Orders Calanoida Cyclopoida Gelyelloida Harpacticoida Misophrioida Monstrilloida Mormonilloida Platycopioida Poecilostomatoida Siphonostomatoida Copepods are a group of small crustaceans found in the sea and nearly every freshwater habitat. ... Any of various small, shrimplike, chiefly marine crustaceans of the order Mysidacea, the females of which carry their eggs in a pouch beneath the thorax. ... Families Euphausiidae Euphausia Dana, 1852 Meganyctiphanes Holt and W. M. Tattersall, 1905 Nematobrachion Calman, 1905 Nematoscelis G. O. Sars, 1883 Nyctiphanes G. O. Sars, 1883 Pseudeuphausia Hansen, 1910 Stylocheiron G. O. Sars, 1883 Tessarabrachion Hansen, 1911 Thysanoessa Brandt, 1851 Thysanopoda Latreille, 1831 Bentheuphausiidae Bentheuphausia amblyops Krill are shrimp-like marine...


Geographical distribution

Clupea harengus distribution on a NASA SeaWIFS image - the main concentrations are in the North Atlantic at the North Sea
Clupea harengus distribution on a NASA SeaWIFS image - the main concentrations are in the North Atlantic at the North Sea

Atlantic herring can be found on both sides of the Atlantic ocean. they have an extensive range that covers the North Atlantic waters such as the Gulf of Maine, the Gulf of St Lawrence, the Bay of Fundy, the Labrador Sea, the Davis Straits, the Beaufort Sea, the Denmark Straits, the Norwegian Sea, the North Sea, the Baltic Sea, the English Channel, the Celtic Sea, and the Bay of Biscay. Although Atlantic herring are found in the northern waters sorrounding the Arctic they are however, not considered to be an Arctic species. Image File history File links Atlantic herring distribution graph uwe kils gfdl self File history Legend: (cur) = this is the current file, (del) = delete this old version, (rev) = revert to this old version. ... Image File history File links Atlantic herring distribution graph uwe kils gfdl self File history Legend: (cur) = this is the current file, (del) = delete this old version, (rev) = revert to this old version. ... This article or section does not cite its references or sources. ... SeaWiFS stands for Sea-viewing Wide Field-of-view Sensor. ... For other uses, see Atlantic (disambiguation) The Atlantic Ocean is Earths second-largest ocean, covering approximately one-fifth of its surface. ... The North Sea is a sea of the Atlantic Ocean, located between the coasts of Norway and Denmark in the east, the coast of the British Isles in the west, and the German, Dutch, Belgian and French coasts in the south. ... The Gulf of Saint Lawrence, the worlds largest estuary, is the outlet of North Americas Great Lakes via the Saint Lawrence River into the Atlantic Ocean. ... The Bay of Fundy (French: baie de Fundy) is a bay located on the Atlantic coast of North America, on the northeast end of the Gulf of Maine between the Canadian provinces of New Brunswick and Nova Scotia. ... Labrador Sea is an arm of the North Atlantic Ocean between Labrador and Greenland. ... Approximate area of the Beaufort Sea, and the disputed waters The Beaufort Sea is a large body of water north of The Northwest Territories, the Yukon, and Alaska and west of Canadas arctic islands that is a part of the Arctic Ocean. ... The Denmark Strait is a strait between Greenland and Iceland. ... The Norwegian Sea (Norwegian: Norskehavet) is part of the North Atlantic Ocean northwest of Norway, located between the North Sea (i. ... The North Sea is a sea of the Atlantic Ocean, located between the coasts of Norway and Denmark in the east, the coast of the British Isles in the west, and the German, Dutch, Belgian and French coasts in the south. ... The Baltic Sea is located in Northern Europe, from 53°N to 66°N latitude and from 20°E to 26°E longitude. ... Satellite view of the English Channel The English Channel (French: La Manche, IPA: , the sleeve), also for some time known in England as the British Sea, is a sea of the Atlantic Ocean that separates the island of Great Britain from northern France, and joins the North Sea to the... Coccoliths in the Celtic Sea. ... Map of the Bay of Biscay. ... The red line indicates the 10°C isotherm in July, commonly used to define the Arctic region border The Arctic is the area around the Earths North Pole. ...


Biological specialities

Herring are amongst the most spectacular schoolers ("obligate schooler" under the old definitions), they aggregate together in groups that consist of thousands to hundreds of thousands of individuals these schools traverse the open oceans . A school of herring in general has a very precise arrangement thus allowing the school to maintain a relatively constant cruising speed. Schools that are made up of an individual stock generally travel in a triangular pattern between their spawning grounds e.g. Southern Norway, their feeding grounds (Iceland) and also their nursery grounds (Northern Norway). Such wide triangular journeys are probably important because herring feast efficiently on their own offspring. A school of herring can react very quickly to evade predators; they have excellent hearing. Around SCUBA divers and ROVs they can form a vacuole ("fountain effect"). The phenomenon of schooling is however, far from understood, especially the implications on swimming and feeding-energetics. Many hypotheses have been put forward to explain the function of schooling, such as predator confusion, reduced risk of being found, better orientation, and synchronized hunting. However, schooling can also have some disadvantages such as: oxygen- and food-depletion, excretion buildup in the breathing media. The school-array probably gives advantages in energy saving although this is a highly controversial and much debated field.


Schools of herring can on calm days sometimes be detected at the surface from more than a mile away by the little waves they form, or from a few meters at night when they trigger the bioluminescence of surrounding plankton ("firing"). All underwater recordings show herring constantly cruising with high speeds up to 108 cm per second, and much higher escape speeds. Bioluminescence is the production and emission of light by a living organism as the result of a chemical reaction during which chemical energy is converted to light energy. ... Photomontage of plankton organisms Plankton are drifting organisms that inhabit the water column of oceans, seas, and bodies of fresh water. ...


Habitat requirements

Atlantic herring are in general very tender and fragile fish. They have extraordinarly large and delicate gill surfaces, and upon contact with foreign matter they can lose their large scales. They have retreated from many of the estuaries worldwide due to high pollution content within the water although in some of the estuaries that have been cleaned up herring have been observed returning. The appearance of their larvae is used as bioindicator for cleaner and better oxygenated waters. gills of a Smooth Newt Gills inside of a tuna head In aquatic organisms, gills are a respiratory organ for the extraction of oxygen from water and for the excretion of carbon dioxide. ... Estuaries and coastal waters are among the most productive ecosystems on Earth, providing numerous ecological, economic, cultural, and aesthetic benefits and services. ... Water pollution Pollution is the release of environmental contaminants. ...


Because of their feeding habits, cruising desire, collective behavior and fragility they are only on display in very few aquaria worldwide, this despite their natural abundance in the ocean. Even with the best facilities that these aquaria can offer they appear slim and slow compared to a quivering school in the wild. Aquarium is also the name of the Russian band, which is also spelled Akvarium A 335,000 gallon (1. ...


Life history

Transparent eggs with the eyes visible, one larva hatched. Observe the yolk.
Transparent eggs with the eyes visible, one larva hatched. Observe the yolk.

There is at least one herring stock spawning in any one month of the year, each race having a different spawning time and place (spring, summer, autumn and winter herrings) in 0 to 5 m off Greenland down to 200 m in autumn (bank) herrings of the North Sea. Eggs are laid on the sea bed, on rock, stones, gravel, sand or beds of algae. "...the fish were darting rapidly about, and those who have opportunity to see the fish spawning in more shallow water ... state that both males and females are in constant motion, rubbing against one another and upon the bottom, apparently by pressure aiding in the discharge of the eggs and milt" (Moore at Cross Island, Maine). Image File history File links larvae herring image uwe kils gfdl self File history Legend: (cur) = this is the current file, (del) = delete this old version, (rev) = revert to this old version. ... Image File history File links larvae herring image uwe kils gfdl self File history Legend: (cur) = this is the current file, (del) = delete this old version, (rev) = revert to this old version. ... The egg yolk is the yellow inside an egg. ...

Freshly hatched larvae in a drop of water besides a match to demonstrate how tiny the larvae are. The black eyes and the yolk are visible.
Freshly hatched larvae in a drop of water besides a match to demonstrate how tiny the larvae are. The black eyes and the yolk are visible.

A female herring may deposit from 20000 up to 40000 eggs, according to her age and size, averaging about 30000. In sexually mature herrings, the genital organs are so large just before spawning commences that they make up about one-fifth of the total weight of the fish. Image File history File links Download high resolution version (1062x801, 195 KB)herring larva in drop of water image uwe kils gfdl File history Legend: (cur) = this is the current file, (del) = delete this old version, (rev) = revert to this old version. ... Image File history File links Download high resolution version (1062x801, 195 KB)herring larva in drop of water image uwe kils gfdl File history Legend: (cur) = this is the current file, (del) = delete this old version, (rev) = revert to this old version. ... The egg yolk is the yellow inside an egg. ...


The eggs sink to the bottom, where they stick in layers or clumps to gravel, seaweeds or stones, by means of their coating mucus, or to any other objects on which they chance to settle.

Juvenile herring. Length ca. 38 mm, ca. 3 months old (still transparent). Visible are the otoliths, the gut, the silvery swimbladder and the heart. Click twice into the image for high resolution.
Juvenile herring. Length ca. 38 mm, ca. 3 months old (still transparent). Visible are the otoliths, the gut, the silvery swimbladder and the heart. Click twice into the image for high resolution.

If the layers get too thick they suffer from oxygen depletion and often die, entangled in a maze of fucus. They need a fair amount of water microturbulence, generally provided by wave action or coastal currents. Survival is highest in crevices and behind solid structures, because many predators feast on openly disposed eggs. The individual eggs are 1 to 1.4 mm in diameter, depending on the size of the parent fish and also on the local race. Incubation time is about 40 days at 3°C (38 F), 15 days at 7°C (45 F), 11 days at 10°C (50 F), they die at temperatures above 19°C (68 F). Image File history File links Download high resolution version (2339x1453, 405 KB)herring juvenile image uwe kils gfdl self File history Legend: (cur) = this is the current file, (del) = delete this old version, (rev) = revert to this old version. ... Image File history File links Download high resolution version (2339x1453, 405 KB)herring juvenile image uwe kils gfdl self File history Legend: (cur) = this is the current file, (del) = delete this old version, (rev) = revert to this old version. ... An otolith, or otoconium is a [[[calcium]] carbonate structure in the saccule or utricle of the inner ear (hence the name otolith, or ear-stone). It can be used for age determination, showing rings of different growth summer/winter, like tree rings. ... The gas bladder (also fish maw, less accurately swim bladder or air bladder) is an internal organ that contributes to the ability of a fish to control its buoyancy, and thus to stay at the current water depth, ascend, or descend without having to waste energy in swimming. ... Species F. serratus F. vesiculosus Fucus is a genus of seaweed that lives in the intertidal zones of rocky shores. ...

Very young larvae imaged in situ in the typical oblique swimming position. The animal in the upper right is in the classical S-shape of the beginning phase of an attack of probably a copepod. The remains of the yolk and the long gut are very well visible in the transparent animal in the middle.
Very young larvae imaged in situ in the typical oblique swimming position. The animal in the upper right is in the classical S-shape of the beginning phase of an attack of probably a copepod. The remains of the yolk and the long gut are very well visible in the transparent animal in the middle.

The larvae are 5 to 6 mm long at hatching, with a small yolk sac that is absorbed by the time a length of 10 mm is reached. Only the eyes are well pigmented (a camera works only with a black housing) the rest of the body is as transparent as possible, and virtually invisible under water and natural luminance conditions. Image File history File links Download high resolution version (883x697, 145 KB)herring larvae in situ image scanned with an ecoSCOPE uwe kils gfdl self File history Legend: (cur) = this is the current file, (del) = delete this old version, (rev) = revert to this old version. ... Image File history File links Download high resolution version (883x697, 145 KB)herring larvae in situ image scanned with an ecoSCOPE uwe kils gfdl self File history Legend: (cur) = this is the current file, (del) = delete this old version, (rev) = revert to this old version. ... Orders Calanoida Cyclopoida Gelyelloida Harpacticoida Misophrioida Monstrilloida Mormonilloida Platycopioida Poecilostomatoida Siphonostomatoida Copepods are a group of small crustaceans found in the sea and nearly every freshwater habitat. ... The egg yolk is the yellow inside an egg. ...


The dorsal fin is formed at 15 to 17 mm, the anal fin at about 30 mm - the ventral fins are visible and the tail becomes well forked at 30 to 35 mm - at about 40 mm the little fish begins to look like a herring.


Larvae diagnostics: The larvae of the herring family are very slender and can easily be distinguished from all other young fish of their distribution range of similar form by the location of the vent, which is so far back that it lies close to the base of the tail. But it requires critical examination to distinguish several clupeoids one from another in their early stages, especially herring from sprats.


At the age of one year they are about 100 mm long, first spawning at 3 years.


Schooling

Atlantic herring are world famous for their huge schools, often numbering in the hundreds of thousands or even millions. One recorded enormous school covered 4 sq. km. in area and reportedly had more than 4 billion fish.

school of juvenile herring close to the surface
school of juvenile herring close to the surface
Underwater video (looping) of a school on its migration to their spawning grounds in the Baltic Sea. With such high speed they can migrate over thousands of kilometers. In the North Atlantic they cruise between Norway and Greenland every year.
Underwater video (looping) of a school on its migration to their spawning grounds in the Baltic Sea. With such high speed they can migrate over thousands of kilometers. In the North Atlantic they cruise between Norway and Greenland every year.


herring school - image Uwe Kils GFDL larger image on http://www. ... herring school - image Uwe Kils GFDL larger image on http://www. ... herring school - (Video by Uwe Kils, free in sense of Wikipedia:Copyrights) File history Legend: (cur) = this is the current file, (del) = delete this old version, (rev) = revert to this old version. ... herring school - (Video by Uwe Kils, free in sense of Wikipedia:Copyrights) File history Legend: (cur) = this is the current file, (del) = delete this old version, (rev) = revert to this old version. ... The Baltic Sea is located in Northern Europe, from 53°N to 66°N latitude and from 20°E to 26°E longitude. ... For other uses, see Atlantic (disambiguation) The Atlantic Ocean is Earths second-largest ocean, covering approximately one-fifth of its surface. ...


Feeding

Slow motion macrophotography video (50% timelag, looping, each image shifted to compensate the rolling microturbulences from the waves) of feeding juvenile herring (38 mm) on copepods - the fish approach from below and catch each copepod individually. In the middle of the image a copepod escapes successfully to the left. Scanned with the ecoSCOPE
Slow motion macrophotography video (50% timelag, looping, each image shifted to compensate the rolling microturbulences from the waves) of feeding juvenile herring (38 mm) on copepods - the fish approach from below and catch each copepod individually. In the middle of the image a copepod escapes successfully to the left. Scanned with the ecoSCOPE
In this sequence a herring attacks four times in a row (50% timelag, looping, each image shifted to compensate the rolling microturbulences from the waves). In the third attack the copepod is visible between the wide opened sides of the mouth. The opercula are spread wide open to compensate the pressure wave which would alert the copepod to trigger a jump.
In this sequence a herring attacks four times in a row (50% timelag, looping, each image shifted to compensate the rolling microturbulences from the waves). In the third attack the copepod is visible between the wide opened sides of the mouth. The opercula are spread wide open to compensate the pressure wave which would alert the copepod to trigger a jump.

Herring is a pelagic feeder - their prey consists of copepods, amphipods, larval snails, diatoms (only herring larvae below 20 mm), peridinians, molluscan larvae, fish eggs, euphausids, mysids, small fishes, herring larvae, menhaden larvae, pteropods, annelids, tintinnids (only herring larvae below 45 mm), Haplosphaera, Calanus, Pseudocalanus, Acartia, Hyperia, Centropages, Temora, Meganyctiphanes norvegica. slow motion video (50%) of feeding juvenile herring on copepods produced with ecoSCOPE - image Uwe Kils File history Legend: (cur) = this is the current file, (del) = delete this old version, (rev) = revert to this old version. ... slow motion video (50%) of feeding juvenile herring on copepods produced with ecoSCOPE - image Uwe Kils File history Legend: (cur) = this is the current file, (del) = delete this old version, (rev) = revert to this old version. ... Orders Calanoida Cyclopoida Gelyelloida Harpacticoida Misophrioida Monstrilloida Mormonilloida Platycopioida Poecilostomatoida Siphonostomatoida Copepods are a group of small crustaceans found in the sea and nearly every freshwater habitat. ... This article needs cleanup. ... video herring - image Uwe Kils GFDL - produced with ecoSCOPE File history Legend: (cur) = this is the current file, (del) = delete this old version, (rev) = revert to this old version. ... video herring - image Uwe Kils GFDL - produced with ecoSCOPE File history Legend: (cur) = this is the current file, (del) = delete this old version, (rev) = revert to this old version. ... Orders Calanoida Cyclopoida Gelyelloida Harpacticoida Misophrioida Monstrilloida Mormonilloida Platycopioida Poecilostomatoida Siphonostomatoida Copepods are a group of small crustaceans found in the sea and nearly every freshwater habitat. ... The operculum in fish is the hard bony flap covering and protecting the gills of Bony fish. ... Orders Calanoida Cyclopoida Gelyelloida Harpacticoida Misophrioida Monstrilloida Mormonilloida Platycopioida Poecilostomatoida Siphonostomatoida Copepods are a group of small crustaceans found in the sea and nearly every freshwater habitat. ... Families about 200 partial list Alpheidae Ampeliscidae Amphilochidae Ampithoidae Anisogammaridae Aoridae Artesiidae Bogideillidae Bosminidae Caprellidae Corophiidae Crangonyctidae Eusiridae Gammaridae Hadziidae Haustoriidae Iphimediidae Ischyroceridae Leucothoidae Liljeborgiidae Lysianassidae Melitidae Phoxocephalidae Sebidae Talitridae Amphipoda (amphipods) include about 4600 different species of small, shrimp_like crustaceans. ... The name snail applies to most members of the molluscan class Gastropoda that have coiled shells. ... Diatoms (Gr. ... Families Euphausiidae Euphausia Dana, 1852 Meganyctiphanes Holt and W. M. Tattersall, 1905 Nematobrachion Calman, 1905 Nematoscelis G. O. Sars, 1883 Nyctiphanes G. O. Sars, 1883 Pseudeuphausia Hansen, 1910 Stylocheiron G. O. Sars, 1883 Tessarabrachion Hansen, 1911 Thysanoessa Brandt, 1851 Thysanopoda Latreille, 1831 Bentheuphausiidae Bentheuphausia amblyops Krill are shrimp-like marine... Any of various small, shrimplike, chiefly marine crustaceans of the order Mysidacea, the females of which carry their eggs in a pouch beneath the thorax. ... Families Limacinidae Cavoliniidae Clioidae Creseidae Cuvierinidae Praecuvierinidae Peraclididae Cymbuliidae Desmopteridae Sea butterflies, or flapping snails, are holoplanktonic mollusks (Mollusca, Gasteropoda), belonging to the suborder Thecosomata (Blainville, 1824). ... Classes and subclasses Class Polychaeta      (paraphyletic?) Class Clitellata    Oligochaeta-      (Earthworms, etc. ... Tintinnids are ciliates of the marine taxon Tintinnida recognized for their formation of vase-shaped coverings called loricae, which often incorporate minute pieces of rock. ... Binomial name Meganyctiphanes norvegica Northern Krill (Meganyctiphanes norvegica) is a crustacean that lives in the North Atlantic Ocean. ...


Young herring capture copepods predominantly individually ("particulate feeding" or "raptorial feeding") (Kils 1992), a feeding method also used by adult herring on large prey items like euphausids.


If prey concentrations reach very high levels, as in microlayers, at fronts or directly below the surface, herring ram forwards with wide open mouth and far expanded opercula over several feet, then closing and cleaning the gill rakers for a few milliseconds ("sift feeding" or "filter feeding").

Herring ram feeding on a school of copepods. All fish have the opercula wide open all at the same time (the red gills are visible) and the mouth wide open (click to enlarge). The fish swim in a grid with a distance of the jumplength of their prey, as indicated in the animation below.
Herring ram feeding on a school of copepods. All fish have the opercula wide open all at the same time (the red gills are visible) and the mouth wide open (click to enlarge). The fish swim in a grid with a distance of the jumplength of their prey, as indicated in the animation below.
Juvenile herring hunt for the very alert and evasive copepods in synchronization: The copepods can sense with their antennae the pressure-wave of the approaching herring and react with a fast escape jump. The length of the jump is quite similar. The fjish arrange in a grid of this characteristic jumplength. The copepods can dart ca. 80 times before they tire out. It takes 60 milliseconds to spread out the antennae again, and this timeslot is utilized often by the herring to snap finally a copepod. A single juvenile herring would never be able to catch a large copepod ("Synchropredation" - results from in situ videos taken from the ATOLL laboratory).
Juvenile herring hunt for the very alert and evasive copepods in synchronization: The copepods can sense with their antennae the pressure-wave of the approaching herring and react with a fast escape jump. The length of the jump is quite similar. The fjish arrange in a grid of this characteristic jumplength. The copepods can dart ca. 80 times before they tire out. It takes 60 milliseconds to spread out the antennae again, and this timeslot is utilized often by the herring to snap finally a copepod. A single juvenile herring would never be able to catch a large copepod ("Synchropredation" - results from in situ videos taken from the ATOLL laboratory).



Image File history File links herring ram feeding image uwe kils gfdl File history Legend: (cur) = this is the current file, (del) = delete this old version, (rev) = revert to this old version. ... Image File history File links herring ram feeding image uwe kils gfdl File history Legend: (cur) = this is the current file, (del) = delete this old version, (rev) = revert to this old version. ... Orders Calanoida Cyclopoida Gelyelloida Harpacticoida Misophrioida Monstrilloida Mormonilloida Platycopioida Poecilostomatoida Siphonostomatoida Copepods are a group of small crustaceans found in the sea and nearly every freshwater habitat. ... Image File history File links synchropredation animation by uwe kils gfdl self File history Legend: (cur) = this is the current file, (del) = delete this old version, (rev) = revert to this old version. ... Image File history File links synchropredation animation by uwe kils gfdl self File history Legend: (cur) = this is the current file, (del) = delete this old version, (rev) = revert to this old version. ... Orders Calanoida Cyclopoida Gelyelloida Harpacticoida Misophrioida Monstrilloida Mormonilloida Platycopioida Poecilostomatoida Siphonostomatoida Copepods are a group of small crustaceans found in the sea and nearly every freshwater habitat. ... Antennae (singular antenna), are the paired appendages connecting to the first (and in crustaceans also to the second) segment of the head of the members of all subphyla of the arthropods except Chelicerata. ... The ATOLL laboratory, here in front of the Kiel powerplant together with netcages with salmonids The Antarctic Technology Offshore Lagoon Laboratory (ATOLL) was a floating oceanographic laboratory for in situ observation experiments and for field testing new instruments and equipment for polar expeditions. ...


References

The Integrated Taxonomic Information System (ITIS) is a partnership designed to provide consistent and reliable information on the taxonomy of biological species. ... 11 March is the 70th day of the year in the Gregorian Calendar (71st in Leap year). ... 2006 (MMVI) is a common year starting on Sunday of the Gregorian calendar. ...

Further reading

  • Bigelow, H.B., M.G. Bradbury, J.R. Dymond, J.R. Greeley, S.F. Hildebrand, G.W. Mead, R.R. Miller, L.R. Rivas, W.L. Schroeder, R.D. Suttkus and V.D. Vladykov (1963) Fishes of the western North Atlantic. Part three New Haven, Sears Found. Mar. Res., Yale Univ.
  • Eschmeyer, William N., ed. 1998 Catalog of Fishes Special Publication of the Center for Biodiversity Research and Information, no. 1, vol 1-3. California Academy of Sciences. San Francisco, California, USA. 2905. ISBN 0-940228-47-5.
  • Fish, M.P. and W.H. Mowbray (1970) Sounds of Western North Atlantic fishes. A reference file of biological underwater sounds The Johns Hopkins Press, Baltimore.
  • Flower, S.S. (1935) Further notes on the duration of life in animals. I. Fishes: as determined by otolith and scale-readings and direct observations on living individuals Proc. Zool. Soc. London 2:265-304.
  • Food and Agriculture Organization (1992). FAO yearbook 1990. Fishery statistics. Catches and landings FAO Fish. Ser. (38). FAO Stat. Ser. 70:(105):647 p.
  • Joensen, J.S. and Å. Vedel Tåning (1970) Marine and freshwater fishes. Zoology of the Faroes LXII - LXIII, 241 p. Reprinted from,

Jonsson, G. (1992). Islenskir fiskar. Fiolvi, Reykjavik, 568 pp.

  • Kinzer, J. (1983) Aquarium Kiel: Beschreibungen zur Biologie der ausgestellten Tierarten. Institut für Meereskunde an der Universität Kiel. pag. var.
  • Koli, L. (1990) Suomen kalat. [Fishes of Finland] Werner Söderström Osakeyhtiö. Helsinki. 357 p. (in Finnish).
  • Laffaille, P., E. Feunteun and J.C. Lefeuvre (2000) Composition of fish communities in a European macrotidal salt marsh (the Mont Saint-Michel Bay, France) Estuar. Coast. Shelf Sci. 51(4):429-438.
  • Landbrugs -og Fiskeriministeriet. (1995). Fiskeriårbogen 1996 Årbog for den danske fiskerflåde Fiskeriårbogens Forlag ved Iver C. Weilbach & Co A/S, Toldbodgade 35, Postbox 1560, DK-1253 København K, Denmark. p 333-338, 388, 389 (in Danish).
  • Linnaeus, C. (1758) Systema Naturae per Regna Tria Naturae secundum Classes, Ordinus, Genera, Species cum Characteribus, Differentiis Synonymis, Locis 10th ed., Vol. 1. Holmiae Salvii. 824 p.
  • Munroe, Thomas, A. / Collette, Bruce B., and Grace Klein-MacPhee, eds. 2002 Herrings: Family Clupeidae. Bigelow and Schroeder's Fishes of the Gulf of Maine, Third Edition. Smithsonian Institution Press. Washington, DC, USA. 111-160. ISBN 1-56098-951-3.
  • Murdy, Edward O., Ray S. Birdsong, and John A. Musick 1997 Fishes of Chesapeake Bay Smithsonian Institution Press. Washington, DC, USA. xi + 324. ISBN 1-56098-638-7.
  • Muus, B., F. Salomonsen and C. Vibe (1990) Grønlands fauna (Fisk, Fugle, Pattedyr) Gyldendalske Boghandel, Nordisk Forlag A/S København, 464 p. (in Danish).
  • Muus, B.J. and J.G. Nielsen (1999) Sea fish. Scandinavian Fishing Year Book Hedehusene, Denmark. 340 p.

0*Muus, B.J. and P. Dahlström (1974) Collins guide to the sea fishes of Britain and North-Western Europe Collins, London, UK. 244 p.

  • Robins, Richard C., Reeve M. Bailey, Carl E. Bond, James R. Brooker, Ernest A. Lachner, et al. 1991 Common and Scientific Names of Fishes from the United States and Canada, Fifth Edition. American Fisheries Society Special Publication, no. 20. American Fisheries Society. Bethesda, Maryland, USA. 183. ISBN 0-913235-70-9.
  • Robins, Richard C., Reeve M. Bailey, Carl E. Bond, James R. Brooker, Ernest A. Lachner, et al. 1991 Common and Scientific Names of Fishes from the United States and Canada, Fifth Edition. American Fisheries Society Special Publication, no. 20. American Fisheries Society. Bethesda, Maryland, USA. 183. ISBN 0-913235-70-9.
  • Whitehead, P.J.P. (1985). FAO species catalogue. Vol. 7. Clupeoid fishes of the world (suborder Clupeioidei). An annotated and illustrated catalogue of the herrings, sardines, pilchards, sprats, shads, anchovies and wolf-herrings Part 1 - Chirocentridae
  • Whitehead, P.J.P. (1985). FAO species catalogue. Vol. 7. Clupeoid fishes of the world (suborder Clupeioidei). An annotated and illustrated catalogue of the herrings, sardines, pilchards, sprats, shads, anchovies and wolf-herrings Part 1 - Chirocentridae, Clupeidae and Pristigaste FAO Fish. Synop. 125(7/1):1-303.
  • Whitehead, Peter J. P. 1985. Clupeoid Fishes of the World (Suborder Clupeoidei): An Annotated and Illustrated Catalogue of the Herrings, Sardines, Pilchards, Sprats, Shads, Anchovies and Wolf-herrings: Part 1 - Chirocentridae, Clupeidae and Pristigasteridae FAO Fisheries Synopsis, no. 125, vol. 7, pt. 1. Food and Agriculture Organization of the United Nations. Rome, Italy. x + 303. ISBN 92-5-102340-9.

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