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Fox was the third son of Henry Fox, 1st Baron Holland, one of the older generation of self-aggrandizing Whigs.
Fox was also a leader of fashion early on, and after a tour of Europe brought back to London the extravagant male fashions then popular at the French court - frilly lace, brocade, cosmetics, red heels etc. This was the costume of the 'Macaronis', and at nineteen Fox was the acknowledged leader of this group.
Fox was made a junior Lord of the Admiralty by Lord North in 1770, but he resigned in January 1772 in order to vote against the Royal Marriages Act, although he was reappointed to a government post at the Treasury in December.