He is well known for originating the four questions he believed should be asked of any animal behaviour, which were:
Function: how does the behaviour impact on the animal's chances of survival and reproduction?
Causation: what are the stimuli that elicit the response, and how has it been modified by recent learning?
Development: how does the behaviour change with age, and what early experiences are necessary for the behaviour to be shown?
Evolutionary history: how does the behaviour compare with similar behaviour in related species, and how might it have arisen through the process of phylogeny?
They are still considered as the cornerstone of modern ethology.
This is a profound biography of the eminent ethologist and Nobel Laureate Nikolaas Tinbergen (1907-1988).
Tinbergen has been one of my scientific heroes for many years, with his brilliant focus on the four whys, his emphasis on the importance of observation and naturalistic studies, his all-time classic "The Study of Instinct", and his many inspiring field experiments, such as the egg shell removal in fl-headed gulls (an antipredator tactic).
Tinbergen was superb as a communicator, both in print and orally, and he was an excellent photographer and film maker in the area of animal behavior.